:root {
  --paper: #fbf8f0;
  --paper-strong: #fffdf8;
  --pine: #123e2d;
  --pine-deep: #0a2a1e;
  --moss: #778f55;
  --sage: #dce5d5;
  --ink: #17201c;
  --muted: #5c655f;
  --rule: #d7d1c3;
  --accent: #2f6d4e;
  --accent-deep: #23543b;
  --focus: #1468b3;
  --serif: Georgia, 'Times New Roman', serif;
  --sans: Inter, ui-sans-serif, -apple-system, BlinkMacSystemFont, 'Segoe UI', sans-serif;
  --container: min(1120px, calc(100vw - 40px));
  --shadow: 0 18px 50px rgba(15, 38, 28, .12);
}

* { box-sizing: border-box; }
html { scroll-behavior: smooth; }
body {
  margin: 0;
  background: var(--paper);
  color: var(--ink);
  font-family: var(--sans);
  font-size: 16px;
  line-height: 1.6;
  text-rendering: optimizeLegibility;
}
img { display: block; max-width: 100%; }
a { color: inherit; text-underline-offset: .18em; }
button, input, textarea { font: inherit; }
button { color: inherit; }
:focus-visible { outline: 3px solid var(--focus); outline-offset: 3px; }
[hidden] { display: none !important; }
.visually-hidden {
  position: absolute !important;
  width: 1px !important;
  height: 1px !important;
  padding: 0 !important;
  margin: -1px !important;
  overflow: hidden !important;
  clip: rect(0, 0, 0, 0) !important;
  white-space: nowrap !important;
  border: 0 !important;
}

.skip-link {
  position: fixed;
  z-index: 1000;
  left: 16px;
  top: -80px;
  padding: 10px 16px;
  background: var(--paper-strong);
  border: 2px solid var(--pine);
}
.skip-link:focus { top: 16px; }

.site-header {
  position: sticky;
  z-index: 50;
  top: 0;
  border-bottom: 1px solid rgba(18, 62, 45, .16);
  background: rgba(251, 248, 240, .96);
  backdrop-filter: blur(12px);
}
.header-inner {
  width: var(--container);
  min-height: 74px;
  margin: 0 auto;
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 28px;
}
.brand { display: inline-flex; align-items: center; gap: 11px; text-decoration: none; }
.brand img { width: 35px; height: 35px; }
.brand-name { color: var(--pine); font: 700 1.55rem/1 var(--serif); }
.site-nav { display: flex; align-items: center; gap: 30px; }
.site-nav a { position: relative; font-size: .91rem; text-decoration: none; }
.site-nav a:not(.button)::after {
  content: '';
  position: absolute;
  height: 1px;
  inset: auto 0 -5px;
  background: currentColor;
  transform: scaleX(0);
  transform-origin: right;
  transition: transform .2s ease;
}
.site-nav a:hover::after, .site-nav a[aria-current='page']::after {
  transform: scaleX(1);
  transform-origin: left;
}
.menu-button { display: none; border: 0; background: transparent; padding: 8px; }
.menu-button span { display: block; width: 24px; height: 2px; margin: 5px; background: var(--pine); }

.button {
  display: inline-flex;
  min-height: 46px;
  align-items: center;
  justify-content: center;
  gap: 10px;
  padding: 10px 19px;
  border: 1px solid var(--pine);
  border-radius: 3px;
  color: var(--pine);
  font-weight: 700;
  text-decoration: none;
  transition: background .2s ease, color .2s ease, transform .2s ease;
}
.button:hover { transform: translateY(-2px); }
.button-primary { border-color: var(--accent-deep); background: var(--accent-deep); color: #fff; }
.button-primary:hover { background: var(--pine); }
.button-dark { border-color: var(--paper); color: var(--paper); }
.text-link { color: var(--accent-deep); font-weight: 700; text-decoration: none; }
.text-link:hover { text-decoration: underline; }

.container { width: var(--container); margin-inline: auto; }
.section { padding: clamp(62px, 7vw, 92px) 0; }
.section-tight { padding: 54px 0; }
.section-label {
  margin: 0 0 15px;
  color: var(--accent-deep);
  font-size: .75rem;
  font-weight: 800;
  letter-spacing: .13em;
  text-transform: uppercase;
}
h1, h2, h3 { margin: 0; color: var(--pine-deep); font-family: var(--serif); font-weight: 500; line-height: 1.04; }
h1 { font-size: clamp(3rem, 5.5vw, 5.4rem); letter-spacing: -.04em; }
h2 { font-size: clamp(2.15rem, 4vw, 3.45rem); letter-spacing: -.03em; }
h3 { font-size: clamp(1.35rem, 2.2vw, 1.9rem); }
.lede { max-width: 650px; margin: 26px 0 0; color: #35403a; font: 1.22rem/1.55 var(--serif); }
.section-intro { max-width: 650px; margin: 19px 0 0; color: var(--muted); font-size: 1.08rem; }

.hero { overflow: hidden; border-bottom: 1px solid var(--rule); background: var(--paper-strong); }
.hero-grid {
  width: var(--container);
  min-height: min(610px, calc(100vh - 74px));
  margin: 0 auto;
  display: grid;
  grid-template-columns: minmax(0, 1.14fr) minmax(360px, .86fr);
  gap: clamp(42px, 6vw, 82px);
  align-items: center;
}
.hero-copy { z-index: 1; padding: 66px 0; }
.hero-copy h1 { max-width: 690px; font-size: clamp(3.15rem, 4.8vw, 4.9rem); }
.hero-copy p { max-width: 560px; margin: 25px 0 32px; color: #39423d; font-size: 1.13rem; }
.hero-actions { display: flex; flex-wrap: wrap; gap: 14px; }
.hero-media { overflow: hidden; width: 100%; max-width: 450px; aspect-ratio: 4 / 3; justify-self: end; box-shadow: var(--shadow); }
.hero-media > img { width: 100%; height: 100%; object-fit: cover; }

.proof-band { border-bottom: 1px solid var(--rule); background: #f2eee4; }
.proof-grid { display: grid; grid-template-columns: repeat(3, 1fr); }
.proof-item { padding: 26px 34px; border-left: 1px solid var(--rule); }
.proof-item:first-child { border-left: 0; }
.proof-item strong { display: block; color: var(--pine); font: 1.35rem var(--serif); }
.proof-item span { color: var(--muted); font-size: .88rem; }

.home-methods { background: #f2eee4; }
.method-grid { display: grid; grid-template-columns: repeat(3, 1fr); border-block: 1px solid var(--rule); }
.method-grid article { padding: 30px 34px 32px 0; }
.method-grid article + article { padding-left: 34px; border-left: 1px solid var(--rule); }
.method-grid span { color: var(--accent); font: italic 1rem var(--serif); }
.method-grid h3 { margin-top: 24px; font-size: 1.45rem; }
.method-grid p { margin: 12px 0 0; color: var(--muted); }

.species-preview { border-block: 1px solid var(--rule); background: var(--paper-strong); }
.section-heading-row { display: flex; align-items: end; justify-content: space-between; gap: 30px; margin-bottom: 38px; }
.section-heading-row .text-link { white-space: nowrap; }
.species-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 34px 26px; }
.species-card { min-width: 0; }
.species-card > a { display: block; text-decoration: none; }
.hero-media, .species-image, .research-image, .research-hero-photo, .species-hero-image, .gallery figure, .maker-image { position: relative; }
.species-image { aspect-ratio: 4 / 3; overflow: hidden; background: var(--sage); }
.species-image img { width: 100%; height: 100%; object-fit: cover; transition: transform .45s ease; }
.photo-credit { position: absolute; z-index: 3; right: 0; bottom: 0; max-width: 80%; padding: 5px; background: rgba(0, 0, 0, .2); color: #fff; font: 8px/1.25 var(--sans); text-align: right; text-shadow: 1px 1px 10px #000; opacity: .7; overflow-wrap: anywhere; pointer-events: none; }
.species-card > a:hover .species-image img { transform: scale(1.025); }
.species-card h3 { margin-top: 17px; }
.latin { margin: 3px 0 12px; color: var(--muted); font-family: var(--serif); font-style: italic; }
.species-meta { display: flex; flex-wrap: wrap; gap: 10px 18px; padding-top: 12px; border-top: 1px solid var(--rule); color: #455149; font-size: .85rem; }

.research-band { position: relative; overflow: hidden; background: var(--pine); color: var(--paper); }
.research-grid { min-height: 520px; display: grid; grid-template-columns: .9fr 1.1fr; align-items: center; }
.research-copy { z-index: 2; padding: 70px 7vw 70px 0; }
.research-copy h2 { color: var(--paper); }
.research-copy p { max-width: 520px; margin: 24px 0 30px; color: #dce5d5; font: 1.15rem/1.55 var(--serif); }
.research-image { align-self: stretch; min-height: 460px; margin-right: calc((100vw - min(1180px, calc(100vw - 40px))) / -2); }
.research-image img { width: 100%; height: 100%; object-fit: cover; }

.maker-grid { display: grid; grid-template-columns: .8fr 1.2fr; gap: clamp(45px, 8vw, 110px); align-items: center; }
.maker-image { aspect-ratio: 4 / 5; overflow: hidden; }
.maker-image img { width: 100%; height: 100%; object-fit: cover; }
.maker-copy p { max-width: 620px; color: var(--muted); }
.about-maker-grid { grid-template-columns: 240px minmax(0, 1fr); align-items: start; }
.about-maker-grid .maker-image { width: 100%; max-width: 240px; }

.safety-band { border-block: 1px solid #d8cdbb; background: #efe8d9; }
.safety-inner { display: grid; grid-template-columns: 80px 1fr auto; align-items: center; gap: 28px; }
.safety-mark { width: 68px; height: 68px; display: grid; place-items: center; border: 2px solid var(--pine); border-radius: 50%; color: var(--pine); font: 2rem var(--serif); }
.safety-band h2 { font-size: 2rem; }
.safety-band p { margin: 6px 0 0; color: #4c514d; }

.site-footer { background: var(--pine-deep); color: #dbe4db; }
.footer-grid { display: grid; grid-template-columns: 1.45fr repeat(3, 1fr); gap: 45px; padding: 64px 0 48px; }
.footer-brand .brand-name { color: #fff; }
.footer-brand .brand img { width: 42px; height: 42px; }
.footer-brand p { max-width: 300px; color: #aebfb4; font-size: .9rem; }
.footer-brand .also-by { margin-top: 20px; }
.footer-brand .also-by a { color: #fff; }
.footer-column h3 { margin-bottom: 17px; color: #fff; font: 700 .75rem var(--sans); letter-spacing: .12em; text-transform: uppercase; }
.footer-column a { display: block; margin: 9px 0; color: #dbe4db; font-size: .9rem; text-decoration: none; }
.footer-column a:hover { text-decoration: underline; }
.footer-bottom { display: flex; justify-content: space-between; gap: 20px; padding: 20px 0 30px; border-top: 1px solid rgba(255,255,255,.15); color: #9fb3a7; font-size: .82rem; }

.page-hero { padding: clamp(64px, 8vw, 102px) 0 62px; border-bottom: 1px solid var(--rule); background: var(--paper-strong); }
.page-hero h1 { max-width: 930px; font-size: clamp(3.1rem, 5.2vw, 4.8rem); }
.page-hero .lede { max-width: 740px; }
.research-page-hero { overflow: hidden; padding: 0; }
.research-hero-grid { min-height: 530px; display: grid; grid-template-columns: minmax(0, 1.08fr) minmax(390px, .92fr); gap: clamp(42px, 6vw, 78px); align-items: center; }
.research-hero-copy { display: flex; flex-direction: column; justify-content: center; padding: 72px 7vw 72px 0; }
.research-hero-copy h1 { font-size: clamp(3rem, 4vw, 4rem); }
.research-hero-photo { overflow: hidden; min-height: 0; aspect-ratio: 4 / 3; box-shadow: var(--shadow); }
.research-hero-photo img { width: 100%; height: 100%; object-fit: cover; }
.breadcrumbs { display: flex; flex-wrap: wrap; gap: 8px; margin: 0 0 30px; color: var(--muted); font-size: .82rem; }
.breadcrumbs a { color: var(--pine); }

.guide-controls { position: sticky; z-index: 20; top: 74px; border-bottom: 1px solid var(--rule); background: rgba(251, 248, 240, .96); backdrop-filter: blur(12px); }
.search-row { display: grid; grid-template-columns: 1fr auto; gap: 24px; padding: 24px 0 18px; align-items: center; }
.guide-search { width: 100%; min-height: 54px; padding: 12px 18px; border: 1px solid #8c948e; border-radius: 3px; background: #fff; color: var(--ink); }
.result-count { color: var(--muted); font-size: .9rem; white-space: nowrap; }
.filter-row { display: flex; gap: 8px; overflow-x: auto; padding: 0 0 18px; scrollbar-width: thin; }
.filter-button { min-height: 40px; padding: 8px 16px; border: 1px solid var(--rule); border-radius: 3px; background: transparent; white-space: nowrap; cursor: pointer; }
.filter-button[aria-pressed='true'] { border-color: var(--pine); background: var(--pine); color: #fff; }
.guide-grid { grid-template-columns: repeat(4, 1fr); }
.guide-results { padding-top: clamp(28px, 4vw, 48px); }
.guide-grid .species-card { padding-bottom: 20px; border-bottom: 1px solid var(--rule); }
.guide-grid .species-card h3 { font-size: 1.45rem; }
.load-more-row { display: flex; justify-content: center; padding-top: 48px; }
.guide-note { display: grid; grid-template-columns: 28px 1fr; gap: 14px; margin-bottom: 38px; padding: 20px; border-left: 3px solid var(--accent); background: #f1eadc; }

.species-hero { width: var(--container); min-height: 500px; margin: 0 auto; display: grid; grid-template-columns: minmax(0, 1fr) minmax(360px, .78fr); gap: clamp(45px, 7vw, 90px); align-items: center; background: var(--paper-strong); }
.species-hero-copy { display: flex; flex-direction: column; justify-content: center; padding: 60px 0; }
.species-hero h1 { max-width: 680px; font-size: clamp(3rem, 4.8vw, 4.7rem); }
.species-hero .latin { margin-top: 14px; font-size: 1.35rem; }
.species-hero-image { width: 100%; max-width: 480px; overflow: hidden; aspect-ratio: 4 / 3; box-shadow: var(--shadow); }
.species-hero-image img { width: 100%; height: 100%; object-fit: cover; }
.species-body { display: grid; grid-template-columns: minmax(0, 1.25fr) minmax(260px, .75fr); gap: clamp(55px, 8vw, 110px); }
.species-copy h2 { margin: 52px 0 16px; font-size: 2.3rem; }
.species-copy h2:first-child { margin-top: 0; }
.species-copy p { max-width: 740px; }
.fact-list { margin: 0; padding: 0; }
.fact-list div { padding: 17px 0; border-top: 1px solid var(--rule); }
.fact-list div:last-child { border-bottom: 1px solid var(--rule); }
.fact-list dt { color: var(--pine); font-size: .77rem; font-weight: 800; letter-spacing: .08em; text-transform: uppercase; }
.fact-list dd { margin: 5px 0 0; }
.gallery { display: grid; grid-template-columns: repeat(2, 1fr); gap: 24px; margin-top: 30px; }
.gallery figure { margin: 0; }
.gallery img { width: 100%; aspect-ratio: 4 / 3; object-fit: cover; }
.gallery figcaption { margin-top: 8px; color: var(--muted); font-size: .76rem; }
.related-list { display: flex; flex-wrap: wrap; gap: 10px; }
.related-list a { padding: 7px 11px; border: 1px solid var(--rule); text-decoration: none; }

.species-research { padding: clamp(64px, 7vw, 92px) 0; border-block: 1px solid var(--rule); background: #f2eee4; }
.species-research-heading { display: grid; grid-template-columns: minmax(0, 1.2fr) minmax(280px, .8fr); gap: 60px; align-items: end; margin-bottom: 38px; }
.species-research-heading h2 { max-width: 760px; font-size: clamp(2.2rem, 4vw, 3.45rem); }
.species-research-heading > p { max-width: 480px; margin: 0 0 5px; color: var(--muted); }
.observation-grid { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 14px; margin-bottom: 34px; }
.observation-card { padding: 24px; border: 1px solid var(--rule); background: var(--paper-strong); }
.observation-card h3 { margin: 0 0 9px; color: var(--pine); font: 800 .76rem/1.3 var(--sans); letter-spacing: .09em; text-transform: uppercase; }
.observation-card p { margin: 0; color: #39443e; font: 1.02rem/1.55 var(--serif); }
.weather-viz { margin-top: 34px; padding: clamp(22px, 4vw, 38px); border: 1px solid var(--rule); background: var(--paper-strong); }
.weather-viz-heading { display: grid; grid-template-columns: minmax(0, 1fr) minmax(280px, .8fr); gap: 48px; align-items: end; margin-bottom: 25px; }
.weather-viz-heading h3 { font-size: clamp(1.85rem, 3vw, 2.55rem); }
.weather-viz-heading > p { margin: 0 0 4px; color: var(--muted); }
.weather-legend { display: flex; flex-wrap: wrap; gap: 12px 26px; margin-bottom: 26px; color: var(--muted); font-size: .82rem; }
.weather-legend span { display: inline-flex; align-items: center; gap: 8px; }
.weather-legend i { width: 16px; height: 16px; border: 1px solid rgba(18, 62, 45, .16); }
.legend-positive { background: rgba(35, 84, 59, .78); }
.legend-negative { background: rgba(168, 116, 53, .78); }
.legend-uncertain { border: 2px solid #8b938e !important; border-radius: 50%; background: var(--paper-strong); }
.confidence-key i { position: relative; height: 2px; border: 0; background: #8b938e; }
.confidence-key i::after { content: ''; position: absolute; inset: -4px auto auto 6px; width: 3px; height: 10px; background: #8b938e; }
.weather-lines { border-top: 1px solid var(--rule); }
.weather-line-row { display: grid; grid-template-columns: 230px minmax(0, 1fr); gap: 22px; align-items: center; padding: 22px 0; border-bottom: 1px solid var(--rule); }
.weather-line-copy h4 { margin: 0 0 8px; color: var(--pine); font-size: 1.35rem; }
.weather-line-copy p { margin: 0; color: var(--muted); font-size: .87rem; line-height: 1.45; }
.weather-line-scroll { min-width: 0; overflow-x: auto; }
.weather-line-chart { display: block; width: 100%; min-width: 620px; height: auto; }
.weather-zone-up { fill: #edf3ee; }
.weather-zone-down { fill: #f5eee4; }
.weather-zero { stroke: #68736c; stroke-width: 1.2; }
.weather-trend { fill: none; stroke: #768079; stroke-width: 1.5; stroke-linejoin: round; stroke-linecap: round; }
.weather-ci, .weather-ci-cap { stroke: #7d867f; stroke-width: 1.2; }
.weather-mark circle { stroke-width: 2; }
.weather-mark-increase circle { fill: var(--accent-deep); stroke: var(--accent-deep); }
.weather-mark-decrease circle { fill: #a87435; stroke: #a87435; }
.weather-mark-uncertain circle { fill: var(--paper-strong); stroke: #7d867f; }
.weather-mark-uncertain .weather-ci, .weather-mark-uncertain .weather-ci-cap { stroke: #a9afa9; }
.weather-y-label, .weather-x-label, .weather-axis-title, .weather-na { fill: var(--muted); font-family: var(--sans); font-size: 11px; }
.weather-y-label { fill: var(--pine); font-weight: 750; }
.weather-y-label-mid { fill: var(--muted); font-size: 10px; font-weight: 600; }
.weather-x-label { font-size: 10px; font-weight: 700; }
.weather-axis-title { font-size: 9px; }
.weather-na { fill: #8b938e; font-weight: 700; }
.weather-research-note { display: flex; align-items: start; justify-content: space-between; gap: 34px; margin-top: 18px; }
.weather-research-note p { max-width: 700px; margin: 0; color: var(--muted); font-size: .82rem; }
.weather-research-note .text-link { flex: none; font-size: .87rem; }

.article-shell { display: grid; grid-template-columns: 220px minmax(0, 760px); gap: clamp(50px, 8vw, 110px); align-items: start; }
.article-toc { position: sticky; top: 112px; }
.article-toc strong { display: block; margin-bottom: 14px; color: var(--pine); font-size: .75rem; letter-spacing: .12em; text-transform: uppercase; }
.article-toc a { display: block; padding: 7px 0; color: var(--muted); font-size: .85rem; text-decoration: none; }
.article-toc a:hover { color: var(--accent-deep); }
.article-body { font-family: var(--serif); font-size: 1.08rem; line-height: 1.78; }
.article-body h2 { margin: 70px 0 20px; font-size: clamp(2.1rem, 4vw, 3.3rem); }
.article-body h2:first-child { margin-top: 0; }
.article-body h3 { margin: 38px 0 10px; font-size: 1.55rem; }
.article-body p { margin: 0 0 22px; }
.article-body .pullquote { margin: 52px 0; padding: 30px 0 30px 35px; border-left: 3px solid var(--accent); color: var(--accent-deep); font: italic 1.8rem/1.35 var(--serif); }
.research-facts { display: grid; grid-template-columns: repeat(3, 1fr); border-block: 1px solid var(--rule); }
.research-fact { padding: 30px 24px; border-left: 1px solid var(--rule); }
.research-fact:first-child { border-left: 0; }
.research-fact strong { display: block; color: var(--pine); font: 2.2rem var(--serif); }
.research-fact span { color: var(--muted); font-size: .82rem; }
.support-callout { margin-top: 60px; padding: 34px; border: 1px solid #cbbca4; background: #f3eadc; }
.support-callout strong { display: block; color: var(--accent-deep); font: 1.7rem/1.25 var(--serif); }
.support-callout span { display: block; margin-top: 10px; }
.references { font-family: var(--sans); font-size: .85rem; }

.content-grid { display: grid; grid-template-columns: minmax(0, 1fr) minmax(260px, .55fr); gap: clamp(50px, 9vw, 120px); align-items: start; }
.content-copy { font-family: var(--serif); font-size: 1.08rem; }
.content-copy h2 { margin: 55px 0 18px; font-size: 2.6rem; }
.content-copy h2:first-child { margin-top: 0; }
.content-aside { padding: 28px; border-top: 3px solid var(--accent); background: #f1eadc; }
.privacy-copy ul { padding-left: 1.25em; }
.privacy-copy li { margin-bottom: 10px; }
.privacy-summary { position: sticky; top: 112px; }
.contact-form { display: grid; gap: 22px; }
.form-field { display: grid; gap: 8px; }
.form-field label { color: var(--pine); font: 750 .82rem/1.3 var(--sans); letter-spacing: .04em; }
.form-field input, .form-field textarea {
  width: 100%;
  padding: 12px 14px;
  border: 1px solid #aaa99f;
  border-radius: 3px;
  background: var(--paper-strong);
  color: var(--ink);
}
.form-field textarea { min-height: 180px; resize: vertical; }
.form-field input:focus, .form-field textarea:focus { border-color: var(--focus); outline: 3px solid rgba(20, 104, 179, .2); outline-offset: 1px; }
.contact-form .button { justify-self: start; cursor: pointer; }
.contact-form .button:disabled { cursor: wait; opacity: .68; transform: none; }
.form-status { min-height: 1.6em; margin: -6px 0 0; color: var(--accent-deep); font: 700 .9rem/1.5 var(--sans); }
.app-screen-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 26px; margin-top: 45px; }
.app-screen { padding: 24px; background: var(--pine); }
.app-screen img { max-height: 620px; margin: 0 auto; }

[data-reveal] { transition: transform .3s ease; }

@media (max-width: 900px) {
  .site-nav { position: absolute; inset: 74px 0 auto; display: none; padding: 25px 20px 30px; border-bottom: 1px solid var(--rule); background: var(--paper-strong); flex-direction: column; align-items: stretch; gap: 18px; }
  .site-nav[data-open='true'] { display: flex; }
  .menu-button { display: block; }
  .hero-grid, .research-grid, .maker-grid, .species-hero, .species-body, .content-grid { grid-template-columns: 1fr; }
  .privacy-summary { position: static; }
  .research-hero-grid { grid-template-columns: 1fr; }
  .research-hero-copy { padding: 65px 0 45px; }
  .research-hero-photo { min-height: 420px; margin: 0; }
  .hero-grid { width: auto; gap: 0; }
  .hero-copy { width: var(--container); margin: 0 auto; padding: 70px 0 48px; }
  .hero-media { width: var(--container); max-width: 560px; margin: 0 auto 58px; justify-self: center; }
  .method-grid { grid-template-columns: 1fr; }
  .method-grid article { padding: 24px 0; }
  .method-grid article + article { padding-left: 0; border-top: 1px solid var(--rule); border-left: 0; }
  .species-grid, .guide-grid { grid-template-columns: repeat(2, 1fr); }
  .research-copy { padding: 70px 0 45px; }
  .research-image { min-height: 400px; margin: 0; }
  .maker-image { max-height: 650px; }
  .about-maker-grid .maker-image { max-width: 220px; }
  .safety-inner { grid-template-columns: 60px 1fr; }
  .safety-inner .button { grid-column: 2; justify-self: start; }
  .footer-grid { grid-template-columns: 1.4fr 1fr 1fr; }
  .footer-brand { grid-column: 1 / -1; }
  .species-hero { gap: 0; }
  .species-hero-copy { padding: 52px 0 38px; }
  .species-hero-image { width: min(100%, 560px); max-width: 560px; margin-bottom: 52px; justify-self: start; }
  .species-research-heading { grid-template-columns: 1fr; gap: 18px; }
  .weather-viz-heading { grid-template-columns: 1fr; gap: 12px; }
  .weather-line-row { grid-template-columns: 1fr; gap: 13px; }
  .article-shell { grid-template-columns: 1fr; }
  .article-toc { position: static; display: flex; gap: 8px 20px; overflow-x: auto; padding-bottom: 15px; }
  .article-toc strong { display: none; }
  .article-toc a { white-space: nowrap; }
  .app-screen-grid { grid-template-columns: 1fr; }
}

@media (max-width: 620px) {
  :root { --container: calc(100vw - 30px); }
  .header-inner { min-height: 66px; }
  .site-nav { inset-block-start: 66px; }
  .site-nav .button { width: 100%; }
  .hero-copy { padding-top: 50px; }
  .hero-media { width: var(--container); }
  .proof-grid { grid-template-columns: 1fr; }
  .proof-item, .proof-item:first-child { padding: 20px 0; border-left: 0; border-top: 1px solid var(--rule); }
  .proof-item:first-child { border-top: 0; }
  .section-heading-row { align-items: start; flex-direction: column; }
  .species-grid, .guide-grid { grid-template-columns: 1fr; }
  .research-image { min-height: 330px; }
  .research-hero-photo { min-height: 330px; }
  .maker-image { max-height: 520px; }
  .safety-inner { grid-template-columns: 1fr; }
  .safety-inner .button { grid-column: auto; }
  .footer-grid { grid-template-columns: 1fr 1fr; gap: 35px; }
  .footer-brand { grid-column: 1 / -1; }
  .footer-bottom { flex-direction: column; }
  .guide-controls { top: 66px; }
  .search-row { grid-template-columns: 1fr; gap: 8px; }
  .species-hero { display: flex; flex-direction: column-reverse; }
  .species-hero-image { width: 100%; min-height: 0; margin-bottom: 0; }
  .gallery { grid-template-columns: 1fr; }
  .observation-grid { grid-template-columns: 1fr; }
  .weather-viz { margin-inline: 0; }
  .weather-research-note { flex-direction: column; gap: 14px; }
  .research-facts { grid-template-columns: 1fr; }
  .research-fact, .research-fact:first-child { border-left: 0; border-top: 1px solid var(--rule); }
  .research-fact:first-child { border-top: 0; }
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { animation-duration: .01ms !important; transition-duration: .01ms !important; }
  [data-reveal] { opacity: 1; transform: none; }
}